salmon grizz


Suraci, J. P., Clinchy, M., Mugerwa, B., Delsey, M., Macdonald, D. W., Smith, J. A., Wilmers, C. C., & Zanette, L. Y. 2016. A new automated behavioural response system to integrate playback experiments into camera trap studies. Methods in Ecology and Evolution. (PDF)

Wheat, R.E., Allen, J.M., Miller, S.D.L., Wilmers, C.C. & Levi, T. 2016. Environmental DNA from Residual Saliva for Efficient Noninvasive Genetic Monitoring of Brown Bears (Ursus arctos). Plos One 11:e0165259. (PDF)

Wilmers, C.C. & Schmitz, O.J. 2016. Effects of gray wolf-induced trophic cascades on ecosystem carbon cycling. Ecosphere 7:1-10. (PDF)

Allen, M.L., Gunther, M.S. & Wilmers, C.C. 2017. The scent of your enemy is my friend? The acquisition of large carnivore scent by a smaller carnivore. Journal of Ethology. (PDF)

Allen, M.L., Wilmers, C.C., Elbroch, L.M., Golla, J.M. & Wittmer, H.U. 2016. The importance of motivation, weapons and foul odors in driving encounter competition in carnivores. Ecology 97: 1905-1912. (PDF)

Wheat, R.E. & Wilmers, C.C. 2016. Habituation reverses fear-based ecologcial effects in brown bears (Ursus arctos). Ecosphere 7:1-11. (PDF)

Smith, J.A., Wang, Y. & Wilmers, C.C. 2016. Spatial characteristics of residential development shift large carnivore prey habits. Journal of Wildlife Management.  (PDF)

Allen, M.L., Yovovich, V. & Wilmers, C.C. 2016. Evaluating the responses of a territorial solitary carnivore to potential mates and competitors. Scientific Reports 6:27257. (PDF)

Gray, M., Wilmers, C.C., Reed, S.E. & Merenlender, A.M. 2016. Landscape feature-based permeability models relate to puma occurrence. Landscape and Urban Planning 147: 50-58. (PDF)


Allen, M.L., Wittmer, H.U., Houghtaling, P., Smith, J., Elbroch, L.M. & Wilmers, C.C. 2015. The Role of Scent Marking in Mate Selection by Female Pumas (Puma concolor). Plos One: 1-12. (PDF)

Levi, T., Wheat, R.E., Allen, J.M. & Wilmers, C.C. 2015. Differential use of salmon by vertebrate consumers: implications for conservation. PeerJ 3:e1157. (PDF)

Wilmers, C.C., Nickel, B., Bryce, C.M., Smith, J.A., Wheat, R.E. & Yovovich, V. 2015. The golden age of biologging:  how animal-borne sensors are advancing the frontiers of ecology. Ecology 96: 1741-1753. (PDF)

Wang, Y., Allen, M.L. & Wilmers, C.C. 2015. Mesopredator spatial and temporal responses to large predators and human development in the Santa Cruz Mountains of California. Biological Conservation 190: 23-33. (PDF)

Allen, M.L., Elbroch, L.M., Wilmers, C.C. & Wittmer, H.U. 2015. The comparative effects of large carnivores on the acquisition of carrion by scavengers. American Naturalist 185: 822-833. (PDF)

Smith, J.A., Wang, Y. & Wilmers, C.C. 2015. Top carnivores increase their kill rates on prey as a response to human-induced fear Proceedings of the Royal Society B 282. (PDF)

Levi, T., Kilpatrick, A.M., Barfield, M., Hold, R.D., Mangel, M. & Wilmers, C.C. 2015. Threshold levels of generalist predation determine consumer response to resource pulses. Oikos. (PDF)

Wang, Y., Nickel, B., Rutishauser, M., Bryce, C., Williams, T.M., Elkaim, G. & Wilmers, C.C. 2015. Movement, resting, and attack behaviors of wild pumas are revealed by tri-axial accelerometer measurements. Movement Ecology 3:2. (PDF)

Allen, M.L., Wallace, C.F. & Wilmers, C.C. 2015. Patterns in bobcat (Lynx rufus) scent marking and communication behaviors. Journal of Ethology 33: 9-14. (PDF)


Williams, T.M., Wolfe, L., Davis, T., Kendall, T., Richter, B., Wang, Y., Bryce, C., Elkaim, G.H. & Wilmers, C.C. 2014. Instantaneous energetics of puma kills reveal advantage of felid sneak attacks. Science 81: 81-85. (PDF)

Tallis, H. et al. 2014. A call for inclusive conservation. Nature 515: 27-28. (PDF)

Schmitt, M.H., Stears, K., Wilmers, C.C. & Shrader, A.M. 2014. Determining the relative importance of dilution and detection for zebra foraging in mixed-species herds. Animal Behaviour 96: 151-158. (PDF)

Allen, M.L., Elbroch, M.L., Wilmers, C.C. & Wittmer, H.U. 2014. Trophic Facilitation or Limitation? Comparative Effects of Pumas and Black Bears on the Scavenger Community. Plos One 9. (PDF)

Allen, M.L., Wittmer, H.U. & Wilmers, C.C. 2014. Puma scrape and communication behaviors: understanding functional use and variation by sex and age. Behaviour 151: 819-840. (PDF)

Ripple, W.J., Estes, J.A., Beschta, R.L., Wilmers, C.C., Ritchie, E.G., Hebblewhite, M., Berger, J., Elmhagen, B., Letnic, M., Nelson, M.P., Schmitz, O.J., Smith, D.W., Wallach, A.D. & Wirsing, A.J. 2014. Status and ecological effects of the world’s largest carnivores. Science 343:151. (PDF, Supplemental)

Schmitz, O.J., Raymond, P.A., Estes, J.A., Kurz, W.A., Holtgrieve, G.W., Ritchie, M.E., Schindler, D.E., Spivak, A.C., Wilson, R.W., Bradford, M.A., Christensen, V., Deegan, L., Smetacek, V., Vanni, M.J. & Wilmers, C.C. 2014. Animating the carbon cycle. Ecosystems 17:344-359. (PDF)


Reid, R.E.B., Greenwald, E.N., Wang, Y. & Wilmers, C.C. 2013. Dietary niche partitioning by sympatric Peromyscus boylii and P.californicus in a mixed evergreen forest. Journal of Mammalogy 94:1248-1257. (PDF)

Wilmers, C.C. & Levi, T. 2013. Do irrigation and predator control reduce the productivity of migratory ungulate herds? Ecology 94: 1264-1270. (PDF)

Wilmers, C.C., Wang, Y. , Nickel,  B., Houghtaling, P., Shakeri, Y., Allen, M. L., Kermish-Wells, J., Yovovich, V., & Williams, T. 2013. Scale dependent behavioral responses to human development by a large predator, the puma. Plos One 8:e60590. (PDF)

Wilmers, C.C., Ram, K., Watson, F.G.R., White, P.J., Smith, D.W. & Levi, T. 2013. Climate and vegetation phenology. Pages 147-163 in P.J. White, R.A. Garrott & G.E. Plumb, editors. Yellowstone’s wildlife in transition. Harvard University Press, Boston. (PDF)

Ripple, W.J., Wirsing, A.J., Wilmers, C.C. & Letnic, M. 2013. Widespread mesopredator effects after wolf extirpation. Biological Conservation 160: 70-79. (PDF)


Wilmers, C.C., Darimont, C.T. & Hebblewhite, M. 2012. Restoring predators as a hedge against climate change. Page 416 in J.F. Brodie, E.S. Post & D.F. Doak, editors. Wildlife conservation in a changing climate. University of Chicago Press, Chicago. (PDF)

Kerby, J.T., Wilmers, C.C. & Post, E. 2012. Climate change, phenology and the nature of consumer-resource interactions: advancing the match/mismatch T. Ohgushi, O.J. Schmitz & R.D. Holt, editors. Trait-mediated indirect interactions: ecological and evolutionary perspectives. Cambridge University Press, Chicago. (PDF)

Wilmers, C.C., Estes, J.A., Laidre, K.L., Edwards, M. & Konar, B. 2012. Do trophic cascades affect the storage and flux of atmospheric carbon? An analysis for sea otters and kelp forests. Frontiers in Ecology and the Environment. (PDF)

Levi, T., Kilpatrick, A.M., Mangel, M. & Wilmers, C.C. 2012. Deer, predators, and the emergence of Lyme disease. Proceedings of the National Academy of Sciences 109: 10942-10947. (PDF)

Levi, T. & Wilmers, C.C. 2012. Wolves-coyotes-foxes: a cascade among carnivores. Ecology 93: 921-929. (PDF)

Levi, T., Darimont, C.T., McDuffee, M., Mangel, M., Paquet, P.C. & Wilmers, C.C. 2012. Using grizzly bears to assess harvest-ecosystem tradeoffs in salmon fisheries. PLoS Biology 10. (PDF)


Rutishauser, M., Petkov, V.V., Boice, J., Obraczka, K., Mantey, P., Williams, T.M. & Wilmers, C.C. 2011. Carnivore: A disruption-tolerant system for studying wildlife. EURASIP Journal on Wireless Communications and Networking. (PDF)

Levi, T., Shepard, G., Ohl-Schacherer, J., Wilmers, C.C., Peres, C. & Yu, D. 2011. Spatial tools for modeling the sustainability of subsistence hunting in tropical forests. Ecological Applications 21: 1802-1818. (PDF)


Darimont, C.T., Bryan, H.M., Carlson, S.M., Hocking, M.D., MacDuffee, M., Paquet, P.C., Price, M.H.H., Reimchen, T.E., Reynolds, J.D. & Wilmers, C.C. 2010. Salmon for terrestrial protected areas. Conservation Letters 3: 379-389. (PDF)


Kinnison, M.T., Palkovacs, E.P., Darimont, C.T., Carlson, S.M., Paquet, P.C. & Wilmers, C.C. 2009. Some cautionary notes on fisheries evolutionary impact assessments. Proceedings of the National Academy of Sciences. 106: E115 (PDF)

Post, E., Brodie, J., Hebblewhite, M., Anders, A.D., Maier, J.A.K. & Wilmers, C.C. 2009. Global population dynamics and hot spots of response to climate change. BioScience. 59: 489-497. (PDF)

Zavaleta, E., Pasari, J., Moore, J., Hernandez, D., Suttle, K. B. & Wilmers, C.C. 2009. Ecosystem responses to community disassembly. The Year in Ecology and Conservation Biology: Ann. N.Y. Acad. Sci. 1162: 311–333. (PDF)

Darimont, C.T., Carlson, S.M., Kinnison, M.T., Paquet, P.C., Reimchen, T.E. & Wilmers, C.C. 2009. Reply to Koons: Harvest related trait change in an increasingly variable world. Proceedings of the National Academy of Sciences 106: E33. (PDF)

Darimont, C.T., Carlson, S.M., Kinnison, M.T., Paquet, P.C., Reimchen, T.E. & Wilmers, C.C. 2009. Human predators outpace other agents of trait change. Proceedings of the National Academy of Sciences 106: 952-954. (PDF)


Post, E., Pederson, C., Wilmers, C.C. & Forchammer, M.C. 2008. Warming, plant phenology and the spatial dimension of trophic mismatch for large herbivores. Proceedings of the Royal Society B. 275: 2005-2013. (PDF)

Post, E., Pederson, C., Wilmers, C.C. & Forchhammer, M.C. 2008. Phenological sequences reveal aggregate life history response to climatic warming. Ecology 89: 363-370. (PDF)


Wilmers, C.C. 2007. Understanding ecosystem robustness. Trends in Ecology and Evolution 22: 504-506. (PDF)

Wilmers, C.C., Post, E. & Hastings A. 2007. The anatomy of predator-prey dynamics in a changing climate. Journal of Animal Ecology 76: 1037-1044. (PDF)

Wilmers, C.C., Post, E. & Hastings A. 2007. A perfect storm: the combined effects on population fluctuations of auto-correlated environmental noise, age structure, and density dependence. American Naturalist 169: 673-683. (PDF)

Getz, W.M., Formann-Roe, S., Cross, P.C., Lyons, A.J., Ryan, S.J., & Wilmers, C.C. 2007. LoCoH: nonparameteric kernel methods for constructing home ranges and utilization distributions. PLoS One 2: e207. (PDF)


Wilmers, C.C., Post, E., Peterson, R.O. & Vucetich, J.A. 2006. Predator disease out-break modulates top-down, bottom-up and climatic effects on herbivore population dynamics. Ecology Letters 9: 383-389. (PDF) See accompanying “Research Highlight” article in Trends in Ecology and Evolution

Wilmers, C.C. & Post, E. 2006. Predicting the influence of wolf-provided carrion on scavenger community dynamics under climate change scenarios. Global Change Biology 12: 403-409. (PDF)


Wilmers, C.C., & Getz, W.M. (2005). Gray wolves as climate change buffers in Yellowstone. PLoS Biology 3: 571-576. (PDF)


Wilmers, C.C., & Getz, W.M. (2004). Simulating the effects of wolf-elk population dynamics on resource flow to scavengers in Yellowstone National Park. Ecological Modelling 177: 193-208. (PDF)

Getz, W.M., & Wilmers, C.C. (2004). A local nearest-neighbor convex-hull construction of home ranges and utilization distributions. Ecography 27: 489-505. (PDF)


Wilmers, C.C., Crabtree, R.L., Smith, D., Murphy, K.M., & Getz, W.M. (2003) Trophic facilitation by introduced top predators: gray wolf subsidies to scavengers in Yellowstone National Park. Journal of Animal Ecology 72: 909-916. (PDF)

Wilmers, C. C., Stahler, D.R., Crabtree, R.L., Smith, D.W.& Getz, W.M. (2003) Resource dispersion and consumer dominance: scavenging at wolf- and hunter-killed carcasses in Greater Yellowstone, USA. Ecology Letters 6: 996-1003. (PDF)


Wilmers, C.C., Sinha, S., & Brede, M. (2002) Examining the effects of species richness on community stability: an assembly model approach. Oikos 99: 363-367. (PDF)

Wilmers, C.C. & Stahler, D.R. (2002) Constraints on active-consumption rates in gray wolves, coyotes and grizzly bears. Canadian Journal of Zoology 80: 1256-1261. (PDF)

Wilmers, C.C. (2002) Diary (on experience researching wolves in Yellowstone NP). London Review of Books 24: 33. (PDF)